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There is no difference—the filters are identical. My poking around the Fleetguard website took me to a page that stated the difference between the FS1268 and the FS19579 is a change to the O-ring that is packaged with the filter. The O-ring that is included with the FS19579 is designed to swell a bit more in the presence of diesel fuel than the O-ring packaged with FS1268.
I have a difficult time navigating fleetguard.com and haven't been able to find the page again (or maybe it no longer exists). The FS 1268 should be fine.
Thomas is %100 correct The FS19579 is the most updated filter, but the only difference is the O'ring. If you have never had a leaking problem then you are good to go with either one. The funny thing is that we here at Cummins Northwest sell the Fs19579 for about $3 less than the fs1268. Go figure.